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About compound leaves?

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Can a compound leaf have a stipule on it's petiole and stipules on it's leaflets?

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  1. Divisions of the lamina (blade)

    Two basic forms of leaves can be described considering the way the blade is divided. A simple leaf has an undivided blade. However, the leaf shape may be formed of lobes, but the gaps between lobes do not reach to the main vein. A compound leaf has a fully subdivided blade, each leaflet of the blade separated along a main or secondary vein. Because each leaflet can appear to be a simple leaf, it is important to recognize where the petiole occurs to identify a compound leaf. Compound leaves are a characteristic of some families of higher plants, such as the Fabaceae. The middle vein of a compound leaf or a frond, when it is present, is called a rachis.

        * Palmately compound leaves have the leaflets radiating from the end of the petiole, like fingers off the palm of a hand, e.g. Cannabis (hemp) and Aesculus (buckeyes).

        * Pinnately compound leaves have the leaflets arranged along the main or mid-vein.

              o odd pinnate: with a terminal leaflet, e.g. Fraxinus (ash).

              o even pinnate: lacking a terminal leaflet, e.g. Swietenia (mahogany).

        * Bipinnately compound leaves are twice divided: the leaflets are arranged along a secondary vein that is one of several branching off the rachis. Each leaflet is called a "pinnule". The pinnules on one secondary vein are called "pinna"; e.g. Albizia (silk tree).

        * trifoliate: a pinnate leaf with just three leaflets, e.g. Trifolium (clover), Laburnum (laburnum).

        * pinnatifid: pinnately dissected to the midrib, but with the leaflets not entirely separate, e.g. Polypodium, some Sorbus (whitebeams).

    Characteristics of the petiole

    The overgrown petioles of Rhubarb (Rheum rhabarbarum) are edible.

    The overgrown petioles of Rhubarb (Rheum rhabarbarum) are edible.

    Petiolated leaves have a petiole. Sessile leaves do not: the blade attaches directly to the stem. In clasping or decurrent leaves, the blade partially or wholly surrounds the stem, often giving the impression that the shoot grows through the leaf. When this is actually the case, the leaves are called "perfoliate", such as in Claytonia perfoliata. In peltate leaves, the petiole attaches to the blade inside from the blade margin.

    In some Acacia species, such as the Koa Tree (Acacia koa), the petioles are expanded or broadened and function like leaf blades; these are called phyllodes. There may or may not be normal pinnate leaves at the tip of the phyllode.

    A stipule, present on the leaves of many dicotyledons, is an appendage on each side at the base of the petiole resembling a small leaf. Stipules may be lasting and not be shed (a stipulate leaf, such as in roses and beans), or be shed as the leaf expands, leaving a stipule scar on the twig (an exstipulate leaf).

        * The situation, arrangement, and structure of the stipules is called the "stipulation".

              o free

              o adnate : fused to the petiole base

              o ochreate : provided with ochrea, or sheath-formed stipules, e.g. rhubarb,

              o encircling the petiole base

              o interpetiolar : between the petioles of two opposite leaves.

              o intrapetiolar : between the petiole and the subtending stem


  2. When a compound leaflet has stipules they are called by another name - stipels or secondary stipules

    http://www.botgard.ucla.edu/html/botanyt...

    http://books.google.com/books?id=wTSO3du...

  3. yes....
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