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I disagree with the statement about the American Lion being larger than the Smilodon Populator.
Do not confuse the Smilodon fatalis, a North American cat, with the South American counterpart, the Smilodon Populator.
Yes, the American Lion was larger than the Smilodon Fatalis ( La Brea Tar Pits Fossil cat ), but it was smaller than the Smilodon Populator, which was larger, heavier, stronger.
It is a huge mistake to think that the Smilodon was a slow animal uncapable of fighting animals and cats its size. This animal evolved large and powerfull, and its smaller cousin, the S Fatalis also did evolve, in an area where other cats also existed, like the Jaguar ( which was larger than today's Jaguar ) the American Lion, the American Cheetah,, a larger animal than the extant Cheetah.
The ferocity and behavioral characteristics that permited the Smilodon to develop, would have assured it a victory if ever confronted to a Lion.
It would be the equivalent to have a Leopard confront a Lion today. Although a strong and agile cat, the Leopard does not have the size and strenght to face off a Lion.
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